Yes it will. You will have to use the Gen 3 recoil spring assembly with the Gen 3 slide. And you will have a gap between the end of the slide and frame, but there are adapters to fill that gap in.
Photo on the left is a Gen slide on a Gen 4/5 frame. Photo on the right is a Gen 3 slide on a Gen 4/5 frame. Notice the gap around the nose of the Gen 3 slide.
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